Apart from horses and dogs, one of the only other species that appears to have a similar brindle pattern is domestic cattle, like the bull above. On the first two dogs, greying has caused the black striping to fade out. Linkage and Segregation Analysis of Black and Brindle Coat Color in Domestic Dogs: http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1931550/ Brindle (kbr) is in the middle of the K locus, recessive to KB but dominant over ky. are sable (AyAy) and tan points (atat). is a brindle. The brindling is still present as normal at skin level, but becomes obscured by long coats. The brindle color, especially on mastiff dog breeds tend to look more monstrous than they will normally look without it. A brindle with stripes that appear to be broken up into shorter stripes and spots may be a merle. This silver brindle Borzoi is most likely an example of a grizzle/domino (Eg) brindle (see below). Here's the quick version! The stunning mixed breed above appears to have some missing stripes on the inside of her front leg. the Skye Terrier just appears to be a muddy grey/brown. This makes it very difficult to test for, and there are currently no commercially available tests that are able to detect brindle. Here are three blue brindles. are sable (AyAy) and tan points (atat). A brindle with stripes that appear to be broken up into shorter stripes and spots may be a merle. Tons of breeds come in brindle, including Akitas, Bull Terriers, Boxers, Boston Terriers, Cardigan Welsh Corgis, Dachshunds, Pit Bulls, and French Bulldogs.. Because the brindle gene allows expression of the A locus, any A locus coat patterns can show brindling on their red sections. The "harlequin" pattern in rabbits, as shown by Wasabi here (submitted by Beth), can look similar to brindle in dogs. Have a look at the Whippet below. As with the Staffordshire Bull Terrier cross shown above (submitted by Ally Stern), the effect can be very subtle. There are many different types of mosaics, but tortoiseshell cats are one type (X-chromosome inactivation means that some cells switch off the black allele and some switch off the red allele - see the "Notes on Mutation etc" page for a full explanation). When the liver and dilution genes are combined, the result is isabella (lilac). A dog that has at least one Em allele on its E locus will have a black mask. Here are three blue brindles. A brindle with stripes that appear to be broken up into shorter stripes and spots may be a merle. This is particularly common in Whippets, Linkage and Segregation Analysis of Black and Brindle Coat Color in Domestic Dogs: http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1931550/, ** Please note that I am not a research scientist, and the information on this page comes from my own knowledge and observation of dogs, observational and testing data provided via e-mail by site visitors, any research papers linked on the page, and the information provided by Dr Sheila M. Schmutz on her excellent website http://homepage.usask.ca/~schmutz/dogcolors.html, For further genetics resources, see the Links page, http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1931550/. Parent club members can be incredibly helpful and knowledgeable, as they have been involved with dogs for many years and will be able to tell you whether the dog looks like their breed, a mixed breed or another breed entirely. Some grizzles can have a greyish cast due to other gene interactions. On some light brindles there may be solid red areas where it appears stripes are "missing". Brindle in dogs is located on the K locus, which is CBD103 (Canine Beta Defensin 103). A brindle is light when the overall appearance is of a red dog and heavy when the overall appearance is of a black dog, although terminology can vary across breeds. The dark (eumelanin) stripes on a brindle can be affected by liver, dilution, greying or merle, and the light/red (phaeomelanin) stripes can be affected by the Intensity locus. ** Please note that I am not a research scientist, and the information on this page comes from my own knowledge and observation of dogs, observational and testing data provided via e-mail by site visitors, any research papers linked on the page, and the information provided by Dr Sheila M. Schmutz on her excellent website http://homepage.usask.ca/~schmutz/dogcolors.html Both the English Mastiff and the American Mastiff can carry the brindle gene. Because the brindle gene allows expression of the A locus, any A locus coat patterns can show brindling on their red sections. are sable (AyAy) and tan points (atat). This is often seen in Japanese Akitas, for example. Brindle is a color pattern, and not a coat color. Variations in Base Colour Have a look at the Whippet below. It is an unstable allele that causes some cells in the body to act as KB (dominant black) and some to act as ky (non-black), in a similar way to a chimaera (fusion of two embryos) or to a tortoiseshell cat (X-chromosome inactivation). Brindle Tan Points and Saddle Markings - Phaeomelanin (the pigment making up the base) can be affected by: Intensity locus. The brindling is still present as normal at skin level, but becomes obscured by long coats. You might be pretty sure, but just need to know the difference between boxers and pit bulls, or your only clue might be the color of your dog's coat. Because a brindle dog will have some cells expressing ky and some expressing KB, this makes testing for brindle extremely difficult. The mechanism does seem to be fairly similar, with some areas of the coat unable to produce eumelanin and some unable to produce phaeomelanin. If a dog is a brindled sable (brindle gene + sable gene), it'll be brindle all over, and on dogs like this masks may be visible. These dogs look very similar to a saddle tan or shaded sable, but the pattern is caused by a different mechanism. Silver Brindle the muzzle and sometimes the ears. The first two photos show Ambrose, owned by Julie, and the third photo was kindly submitted by Abby L A dog that has at least one Em allele on its E locus will have a black mask. There are currently no commercial tests available for the kbr allele, but the allele has been identified by at least one laboratory. Brindle colors range from tawny to reddish to brown to dark black, and there are are many edible items that reflect these colors and make for fun brindle dog names. There are three K locus alleles - KB (dominant black), kbr (brindle) No time to read the whole thing? This is because the pale base can cause the black stripes to appear lighter than they actually are. If a dog is kbrkbr or kbrky then it will display its A locus genes, just like a kyky dog, but any phaeomelanin (red/tan) in the coat will be brindled. the Skye Terrier just appears to be a muddy grey/brown. Sophie, submitted by Helen Lottridge Brindle (kbr) is in the middle of the K locus, recessive to KB but dominant over ky. If you're still unsure about your dog, you can also contact your local veterinarian to get their professional opinion. The "harlequin" pattern in rabbits, as shown by Wasabi here (submitted by Beth), can look similar to brindle in dogs. Because the brindle gene allows expression of the A locus, any A locus coat patterns can show brindling on their red sections. the extent of the brindling on the coat depends on the A locus. the Skye Terrier just appears to be a muddy grey/brown. This is particularly common in Whippets, The brindle pattern is often referred to as ‘tiger-striped’ even though the pattern is more elusive with irregular streaks of color that are darker than the color of the base coat. The base colour on a brindle can vary as much as red can usually vary, from a deep Irish Setter colour to a light cream. "reverse brindle". Dogs with the Eg gene (Salukis, Borzoi, Afghan Hounds) are affected differently by brindle, with the brindling appearing mostly on the dark (eumelanin) areas of the dog. No time to read the whole thing? The overall effect may be similar to how a blue brindle (dd dilution gene) would look. In fact, in horses there is no gene for brindle, and when brindle occurs it is always indicative of a chimaera. However, the light base colour can make the stripes seem greyish (see below). No time to read the whole thing? The visual effect is somewhere inbetween tortoiseshell in cats and dog brindle, as the effect on more heavily marked rabbits is of large stripe-like patches. ** Please note that I am not a research scientist, and the information on this page comes from my own knowledge and observation of dogs, observational and testing data provided via e-mail by site visitors, any research papers linked on the page, and the information provided by Dr Sheila M. Schmutz on her excellent website http://homepage.usask.ca/~schmutz/dogcolors.html These two Greyhounds represent a typical "light" or "red" brindle and a typical "heavy" or "dark" brindle. The "harlequin" pattern in rabbits, as shown by Wasabi here (submitted by Beth), can look similar to brindle in dogs. Look up a comprehensive list of dog breeds with brindle coats. Here's the quick version! Liver and Dilute Brindles These two dogs are brindle with saddle markings(atat plus the RALY saddle tan modifier, see the Agouti page). There are many different types of mosaics, but tortoiseshell cats are one type (X-chromosome inactivation means that some cells switch off the black allele and some switch off the red allele - see the "Notes on Mutation etc" page for a full explanation). Brindle merles are dealt with on the merle page. The Dachshund here has very distinct stripes and is obviously a brindle, but The Brindle Gene On some light brindles there may be solid red areas where it appears stripes are "missing". and isabella stripes if it is both dd and bb. Because a brindle dog will have some cells expressing ky and some expressing KB, this makes testing for brindle extremely difficult. Sometimes dogs with black stripes and a very light cream base (see above) are also referred to as silver brindles. Dogs with the Eg (grizzle/domino) allele seem to show brindle mostly on their darker areas or saddles, as the Saluki puppy above shows. Quick Summary! Learn about all the different types of breeds that can have brindle coats. are sable (AyAy) and tan points (atat). The Whippet is a very light brindle, and the blue colouration is mostly visible on the mask and nose. - Eumelanin (the pigment making up the stripes) can be affected by: merle (and harlequin), liver, dilution, greying, and recessive red. In commercial testing, brindles generally test as KBky. Note the large areas of solid black on the back and brindled tan These two dogs are brindle with saddle markings(atat plus the RALY saddle tan modifier, see the Agouti page). A brindle dog will have liver (brown) stripes if it is bb on the B locus, blue stripes if it is dd on the D locus, A tan pointed dog with the brindle gene will appear black/liver/blue/isabella with brindle points instead of tan. For further genetics resources, see the Links page. Brindle Merles Isabella brindles are extremely difficult to distinguish from a solid isabella as the stripes and the base coat are such a similar colour. Links to studies: Brindle dogs have one or two copies of the brindle gene (from one or both parents). The visual effect is somewhere inbetween tortoiseshell in cats and dog brindle, as the effect on more heavily marked rabbits is of large stripe-like patches. English bulldogs may have a brindle coat. Dogs with the Eg (grizzle/domino) allele seem to show brindle mostly on their darker areas or saddles, as the Saluki puppy above shows. A brindle-colored Frenchie may appear dark or light depending on the hair patterns. the muzzle and sometimes the ears. Brindle dogs will usually test as KBky, and carriers (one dominant black allele, one brindle) cannot be identified without breeding. These two Greyhounds represent a typical "light" or "red" brindle and a typical "heavy" or "dark" brindle. Some grizzles can have a greyish cast due to other gene interactions. Brindle is one of the alleles on the K locus. Next comes the brindle gene, which has the unexpected abbreviation K br. It also appears to have some lighter and darker areas, which also suggest it Here are three blue brindles. Links to studies: The Whippet is a very light brindle, and the blue colouration is mostly visible on the mask and nose. Further Info and Links Dogs with the Eg (grizzle/domino) allele seem to show brindle mostly on their darker areas or saddles, as the Saluki puppy above shows. is a brindle. A light These are all details taken from photographs of brindles with black stripes. Visit the website of the parent club for each breed standard and view the photos of the breed. A dog that has at least one Em allele on its E locus will have a black mask. Brindle (kbr) is in the middle of the K locus, recessive to KB but dominant over ky. Dilution affects phaeomelanin (red) too to some extent, so the base colour Interestingly, a form of mosaicism in humans results in a brindle-like pattern known as Blaschko lines. and heavy when the overall appearance is of a black dog, although terminology can vary across breeds. For further genetics resources, see the Links page. Currently, GSDs have eleven dog kennel club-accepted coat colors. The stunning mixed breed above appears to have some missing stripes on the inside of her front leg. In fact, in horses there is no gene for brindle, and when brindle occurs it is always indicative of a chimaera. Dogs with the Eg gene (Salukis, Borzoi, Afghan Hounds) are affected differently by brindle, with the brindling appearing mostly on the dark (eumelanin) areas of the dog. This is an area of eumelanin covering ** Please note that I am not a research scientist, and the information on this page comes from my own knowledge and observation of dogs, observational and testing data provided via e-mail by site visitors, any research papers linked on the page, and the information provided by Dr Sheila M. Schmutz on her excellent website http://homepage.usask.ca/~schmutz/dogcolors.html Linkage and Segregation Analysis of Black and Brindle Coat Color in Domestic Dogs: http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1931550/ The brindling is still present as normal at skin level, but becomes obscured by long coats. There are many different types of mosaics, but tortoiseshell cats are one type (X-chromosome inactivation means that some cells switch off the black allele and some switch off the red allele - see the "Notes on Mutation etc" page for a full explanation). Sophie, submitted by Helen Lottridge A sable brindle will As urajiro only affects phaeomelanin, the black stripes are unaffected. Crossbreed photo by (unknown) Brindle dogs are not chimaeras, however, but they are mosaics, which is when different cells in the body become genetically distinct during development. On some light brindles there may be solid red areas where it appears stripes are "missing". Liver and Dilute Brindles Color & Marking Description: Color and Markings for your breed. Liver brindles. Apart from horses and dogs, one of the only other species that appears to have a similar brindle pattern is domestic cattle, like the bull above. This means that In the first picture the brindle appears normal, however in the second picture you "reverse brindle". It is an unstable allele that causes some cells in the body to act as KB (dominant black) and some to act as ky (non-black), in a similar way to a chimaera (fusion of two embryos) or to a tortoiseshell cat (X-chromosome inactivation). All of these dogs display black-pigmented dogs with brindled points, with or without white markings. Brindle Saluki puppy photo submitted by Tara Meikle "Missing" Stripes There is one apparent exception to the rule that only the tan areas on a dog show brindle striping. A sable brindle will The thickness of the stripes can vary, so the dog may look predominantly dark or predominantly red. will be relatively light (certainly not a deep red colour, but more likely cream). As with the Staffordshire Bull Terrier cross shown above (submitted by Ally Stern), the effect can be very subtle. occurs almost exclusively on brindles and not on normal solid reds or sables. Linkage and Segregation Analysis of Black and Brindle Coat Color in Domestic Dogs: http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1931550/ Some might also have a small white patch on their chest. This type of striping is sometimes called "black brindle" or A sable brindle will These two dogs are brindle with saddle markings(atat plus the RALY saddle tan modifier, see the Agouti page). A sable brindle will Brindle Tan Points and Saddle Markings It is entirely dominated by KB (so just one KB allele will stop brindle from being expressed), but is dominant over ky, so a brindle dog can have the genotype Pure white, solid blue, merle and brindle are coat colors that some regard as faults and sadly, are slowly coming to extinction. This Glen of Imaal's brindling is very obvious on the short fur on its body, but less obvious on the longer fur on its head. Isabella brindles are extremely difficult to distinguish from a solid isabella as the stripes and the base coat are such a similar colour. A sable brindle will And of course, white markings and ticking can occur on any brindle dog. This silver brindle Borzoi is most likely an example of a grizzle/domino (Eg) brindle (see below). The dark (eumelanin) stripes on a brindle can be affected by liver, dilution, greying or merle, and the light/red (phaeomelanin) stripes can be affected by the Intensity locus. However, the light base colour can make the stripes seem greyish (see below). This is because the pale base can cause the black stripes to appear lighter than they actually are. The Harlequin Coat Pattern. A brindle with stripes that appear to be broken up into shorter stripes and spots may be a merle. Many dogs have a unique coloring called a brindle, which appears as either brown or a dusky tawny color with streaks of other colors mixed in. Here are three blue brindles. There is only a very small bit of tan visible on the hip, and the rest of the The main patterns The Whippet is a very light brindle, and the blue colouration is mostly visible on the mask and nose. Sometimes a brindle dog with black pigment may have what appear to be silver stripes, usually on a cream background (never deep red). The overall effect may be similar to how a blue brindle (dd dilution gene) would look. This is because the pale base can cause the black stripes to appear lighter than they actually are. and isabella stripes if it is both dd and bb. The Neapolitan Mastiff, on the other hand, is a heavy brindle, so the overall appearance is blue. There is one apparent exception to the rule that only the tan areas on a dog show brindle striping. are sable (AyAy) and tan points (atat). The "harlequin" pattern in rabbits, as shown by Wasabi here (submitted by Beth), can look similar to brindle in dogs. The base colour on a brindle can vary as much as red can usually vary, from a deep Irish Setter colour to a light cream. It also appears to have some lighter and darker areas, which also suggest it Brindle is the name given to a coat color pattern characterized by a brown base color, overlain with dark brown to black stripes. As with the Staffordshire Bull Terrier cross shown above (submitted by Ally Stern), the effect can be very subtle. The thickness of the stripes can vary, so the dog may look predominantly dark or predominantly red. As with the Staffordshire Bull Terrier cross shown above (submitted by Ally Stern), the effect can be very subtle. The most popular color? There are many different types of mosaics, but tortoiseshell cats are one type (X-chromosome inactivation means that some cells switch off the black allele and some switch off the red allele - see the "Notes on Mutation etc" page for a full explanation). In fact, in horses there is no gene for brindle, and when brindle occurs it is always indicative of a chimaera. Compare the above factors to the AKC standards to narrow down your list of possible breeds. but the genetic basis is unknown. The main patterns Silver brindle is often caused by the greying gene (G locus), which turns black to grey as the dog ages. Some grizzles can have a greyish cast due to other gene interactions. Brindle Tan Points and Saddle Markings Because a brindle dog will have some cells expressing ky and some expressing KB, this makes testing for brindle extremely difficult. kbr sits between the two. There is one apparent exception to the rule that only the tan areas on a dog show brindle striping. This is the opposite of a regular brindle mastiff appearance. The stunning mixed breed above appears to have some missing stripes on the inside of her front leg. A brindle with stripes that appear to be broken up into shorter stripes and spots may be a merle. RED BRINDLE and WHITE A sable brindle will Some grizzles can have a greyish cast due to other gene interactions. will be relatively light (certainly not a deep red colour, but more likely cream). 2.Brindle Mastiffs are not common. Brindle in dogs is located on the K locus, which is CBD103 (Canine Beta Defensin 103). The K locus determines whether a dog is solid black (K B) or brindle (K br). For further genetics resources, see the Links page. It is an unstable allele that causes some cells in the body to act as KB (dominant black) and some to act as ky (non-black), in a similar way to a chimaera (fusion of two embryos) or to a tortoiseshell cat (X-chromosome inactivation). There is one apparent exception to the rule that only the tan areas on a dog show brindle striping. Here are three blue brindles. Is it a small, medium or large breed? Silver Brindle The stunning mixed breed above appears to have some missing stripes on the inside of her front leg. Isabella brindles are extremely difficult to distinguish from a solid isabella as the stripes and the base coat are such a similar colour. Quick Summary! but the genetic basis is unknown. Even between longhaired breeds, the visibility of brindle can vary. Dogs with the Eg gene (Salukis, Borzoi, Afghan Hounds) are affected differently by brindle, with the brindling appearing mostly on the dark (eumelanin) areas of the dog. Note whether the ears are floppy or upright and the length of the tail. Have a look at the Whippet below. There are many different types of mosaics, but tortoiseshell cats are one type (X-chromosome inactivation means that some cells switch off the black allele and some switch off the red allele - see the "Notes on Mutation etc" page for a full explanation). The harlequin is also a specific color pattern. If they don't have many photos, follow their links to a regional club or a local breeders' website. KB is the top dominant and ky is the bottom recessive. ** Please note that I am not a research scientist, and the information on this page comes from my own knowledge and observation of dogs, observational and testing data provided via e-mail by site visitors, any research papers linked on the page, and the information provided by Dr Sheila M. Schmutz on her excellent website http://homepage.usask.ca/~schmutz/dogcolors.html You may notice that on the Great Dane above the phaeomelanin (red) is deeper on the back Sophie, submitted by Helen Lottridge The coat has a base color with slightly darker striping. Compare these photos to your dog and find your best possible match. Sometimes dogs with black stripes and a very light cream base (see above) are also referred to as silver brindles. The mechanism does seem to be fairly similar, with some areas of the coat unable to produce eumelanin and some unable to produce phaeomelanin. It is an unstable allele that causes some cells in the body to act as KB (dominant black) and some to act as ky (non-black), in a similar way to a chimaera (fusion of two embryos) or to a tortoiseshell cat (X-chromosome inactivation). The main way we can tell this Skye is brindled is simply its muddy-coloured coat - this shade Brindle Merles The stunning mixed breed above appears to have some missing stripes on the inside of her front leg. Sophie, submitted by Helen Lottridge It is an unstable allele that causes some cells in the body to act as KB (dominant black) and some to act as ky (non-black), in a similar way to a chimaera (fusion of two embryos) or to a tortoiseshell cat (X-chromosome inactivation).

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